In Darwin’s Footsteps (Eastern Route)

Explore the incredible biodiversity of the Galápagos Islands on this HX – Hurtigruten Expeditions journey that truly follows in Charles Darwin’s footsteps. Starting with two nights in vibrant Quito, you’ll then fly to the archipelago to cruise among the UNESCO‑protected islands of San Cristóbal, Santa Fe, Santa Cruz, Española, Eden Islet, and North Seymour. Throughout the expedition aboard the intimate MS Santa Cruz II, you’ll encounter iconic wildlife such as giant tortoises, marine iguanas, blue‑footed boobies, and playful sea lions in their natural habitats.

This immersive voyage blends guided shore landings, small‑boat explorations, snorkeling and kayaking opportunities, and enriching insights from knowledgeable Expedition Team members. From beach walks and deep dives into evolutionary history to visits to the Charles Darwin Research Station, every day brings new discovery and wonder. Travel in comfort on an eco‑minded expedition ship designed for up‑close encounters, then return home with stories of one of the planet’s most extraordinary ecosystems.

What’s Included:

Full Inclusion List

Starting from $6,843

per person

Galápagos Islands – In Darwin’s Footsteps (Eastern Route)

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What’s Included:
All Inclusive
Cabin Included
Daily Excursions
Domestic Airfare Included
Expedition Gear
Gratuities Included
Hotel Included

Departure Dates & Pricing:

April 8 – 16, 2026:

  • Starting at $6,843  per person
  • MS Santa Cruz II

April 22 – 30, 2026:

  • Starting at $7,986 per person
  • MS Santa Cruz II

May 6 – 14, 2026:

  • Starting at $7,224 per person
  • MS Santa Cruz II

May 20 – May 28, 2026:

  • Starting at $6,843 per person
  • MS Santa Cruz II

June 3 – 11, 2026:

  • Starting at $6,985 per person
  • MS Santa Cruz II

June 17 – 25, 2026:

  • Starting at $6,985 per person
  • MS Santa Cruz II

July 1– 9, 2026:

  • Starting at $7,605 per person
  • MS Santa Cruz II

July 15 – 23, 2026:

  • Starting at $6,843 per person
  • MS Santa Cruz II

July 29 – August 6, 2026:

  • Starting at $7,605 per person
  • MS Santa Cruz II

August 26 – September 3, 2026:

  • Starting at $6,843 per person
  • MS Santa Cruz II

September 9 – 17, 2026:

  • Starting at $7,224 per person
  • MS Santa Cruz II

September 23 – October 1, 2026:

  • Starting at $7,224 per person
  • MS Santa Cruz II

October 7 – 15, 2026:

  • Starting at $7,224 per person
  • MS Santa Cruz II

October 21 – 29, 2026:

  • Starting at $7,224 per person
  • MS Santa Cruz II

November 4 – 12, 2026:

  • Starting at $7,605 per person
  • MS Santa Cruz II

November 18 – 26, 2026:

  • Starting at $7,224 per person
  • MS Santa Cruz II

December 2 – 10, 2026:

  • Starting at $7,224 per person
  • MS Santa Cruz II

December 16 – 24, 2026:

  • Starting at $7,754 per person
  • MS Santa Cruz II

December 30 – January 7, 2026:

  • Starting at $7,553 per person
  • MS Santa Cruz II

Expires: 2/28/2026 • Supplier Code: HX

*Pricing and availability subject to change until booked. Pricing, unless noted otherwise, is based on double occupancy. Restrictions may apply.

Itinerary:

We’ll help you plan where to go, when to visit and how long to stay.
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Begin your adventure in the vibrant city of Quito

Your journey begins on the Equator in the lush Andean foothills. At 9,350 feet, Quito is the second-highest capital city in the world, and its main draw is the exquisite Old Town which was founded on an ancient Inca settlement. A UNESCO World Heritage site, it’s packed with well-preserved colonial architecture as evidenced in colourful plazas, magnificent churches and handsome monuments.

Settle into your hotel and enjoy an overnight stay.

If you have time, why not come a few days earlier and join one of our pre-voyage options.

Straddle the Equator in the Middle of the World City

Today, enjoy a guided tour of Quito’s remarkable Old Town, one of the largest and best-preserved colonial-era quarters in South America. Wander through cobblestone streets as you admire the architecture and sample delicious chocolate made from the finest Ecuadorian cacao along the way.

Next, you’ll head to the iconic ‘Middle of the World City’, located around 16 miles north of Quito. After lunch, you’ll visit the famous monument that delineates the imaginary line that is the Equator. Have your camera ready to capture the moment you stand with one foot in each hemisphere.

Get ready to explore the spectacular Galápagos Islands

After breakfast, a short flight will take you to Baltra Island, a part of the spectacular Galápagos Archipelago. A transfer will take you straight to the port, where your comfortable expedition ship MS Santa Cruz II awaits you.

Get your first glimpse of the islands’ incredible wildlife on Mosquera Islet, home to one of the largest sea lion colonies in the archipelago. We’ll take our small panga boats ashore for a closer look.

As you stroll across the flat, sandy dunes, look out for blue-footed boobies and Sally Lightfoot crabs, and learn about the marine and terrestrial ecosystems from your naturalist guides.

Meet the famous giant tortoises

Your first stop today is the Cerro Colorado Tortoise Reserve, a breeding center for the endangered giant tortoise. It’s also home to other endemic species, including the San Cristóbal mockingbird and the lava lizard.

The giant tortoise is probably the most iconic of all Galápagos wildlife, with a lifespan of over 100 years. They thrive here, ambling around forests similar to their natural habitat.

Up to August 2024 we’ll call at Cerro Brujo, on the island’s west coast. With its white coral beaches, you’ll literally be walking in Darwin’s footsteps as the naturalist visited here in 1835. There are plenty of sea and shore birds to look out for here, as well as sea lions.

After August 2024* we’ll instead likely explore Punta Pitt in our small panga boats. This is the only site in the Galápagos Islands where you might see three species of boobies. Enjoy a nature walk, explore the coastline by boat, or snorkel and swim in the crystal-clear water.    *Note – this is dependent on local authority restrictions.

Spot wildlife against idyllic backdrops

We start the day with a scenic nature walk on the blissful beaches of Santa Fe, home to the endemic Barrington land iguana and a colony of sea lions.

You can swim or snorkel from our small panga boats, take a kayak for a paddle around the island, or join a trip in our glass-bottomed boat to observe the marine life below. After a delicious lunch prepared by our onboard chefs, we head to South Plaza Island.

Keep an eye out for frigatebirds, swallow-tailed gulls, and Galápagos shearwaters gliding on the breeze. Here, among a dazzling array of flora, land iguanas laze by the shore while marine iguanas slip in and out of the water.

Learn about local conservation efforts on Santa Cruz Island

Today, we pay a visit to the Charles Darwin Research Station in Puerto Ayora.

At this giant tortoise breeding center, you can see recently hatched tortoises and Darwin’s famous finches – the birds he used as a basis for his theory of evolution by natural selection. There’s also an impressive forest of prickly pear cacti that is home to many land birds.

Enjoy a cooler climate over lunch in the lush highlands, where giant tortoises shuffle about munching on grass and wallowing in pools.

You’ll also have time for another activity, such as mountain biking or visiting a sugar plantation.

Lava lizards and incredible birdlife

Your Galápagos Islands cruise reaches Española, the archipelago’s southernmost island. Birdlife here is truly extraordinary. On cruises from April through December, look for Nazca boobies, blue-footed boobies, swallow-tailed gulls, and the mighty waved albatross.

This is also a great chance to spot red Española lava lizards and marine iguanas, as well as see El Soplador, the famous blowhole that blasts water 75 feet into the air.

After lunch on board the ship, we’ll head to the picturesque coral beach of Gardner Bay and nearby Osborn Islet to seek out sea lions, mockingbirds, and finches. If you’re feeling active, you can go snorkeling or kayaking.

Green turtles, sharks, and rare bird colonies

The abundant wildlife-watching opportunities continue at Eden Islet, where we hope to see Galápagos green turtles, rays, and friendly reef sharks. On shore, look for distinctive frigatebirds with their rubbery, red throats.

You can swim or snorkel to explore the reefs. You can also see the reefs from our glass-bottomed boat or by kayak. After lunch, we’ll sail to North Seymour.

The island is a great site for spotting colonies of blue-footed boobies and swallow-tailed gulls. Sea lions and marine iguanas also congregate here, attracted by the bountiful sea life in its waters.

From the Galápagos Islands back to Quito

Your expedition cruise draws to a close as you disembark at Baltra Island. A transfer will take you to the airport in Baltra for your flight to Quito – or if you prefer, to Guayaquil – to to connect to your international flight. To explore more of Ecuador, why not extend your stay and join one of our post-voyage options.

As you leave this fascinating region, you’ll take with you incredible memories of amazing wildlife and wonderful experiences few have the privilege of seeing.

Optional Extras:

It’s a mixture of adventure activities for individuals of all ages.

Galápagos Islands (7)

Mashpi Reserve: A Luxury Rainforest Retreat (Pre)

Day 1: Arrival in Quito

Arrive in Ecuador’s capital, be transferred to your hotel. The rest of the day is yours to explore the city’s historic center. Stroll through its charming streets, visit landmarks such as Plaza Grande, or simply relax and recharge ahead of your adventure.

Day 2: Quito to the Mashpi Reserve

Travel into the Mashpi Reserve, a protected cloud forest in the Chocó hotspot. The three-hour journey from the Andes reveals dramatic shifts in altitude and scenery, descending from 9,200 to 3,300 feet above sea level. On arrival at Mashpi Lodge, enjoy lunch, a conservation briefing, and a guided walk through the flora and fauna along the dense jungle trails. In the evening, listen to fascinating lectures about the unique ecosystem.

Days 3–4: Exploring the Mashpi Reserve

Spend two days discovering the rainforest with your guide. Start at dawn with optional birdwatching, then choose from waterfall hikes, a Sky Bike canopy tour, Dragonfly canopy ride, and visits to the Observation Tower or Life Center. After dark, join a night walk or relax in the lodge’s wellness areas, followed by an Ecuadorian-inspired dinner overlooking the forest below.

Day 5: Return to Quito

Join an early birdwatching walk to see feeding flocks before breakfast. Afterwards, there’s time for a short hike or final activity before you return to Quito. You’ll spend two nights at your hotel, with a city tour scheduled for the following day, before setting off on your Galápagos expedition.

Please note that we may visit sites in a different order.

Galápagos Islands (5)

Jungle Adventure in Ecuador’s Amazon (Post)

Day 1: Arriving at Quito 

Fly from the Galápagos to Quito, where your expedition cruise concludes. We’ll meet you on arrival and transfer you to your airport hotel. For the rest of the day, enjoy some time to relax and unwind before your unforgettable Amazon adventure begins.

Day 2: Rainforest Ecolodge Exploration

After a short, scenic flight over the Andes to Coca, enjoy a two-hour Napo River canoe ride, keeping an eye out for herons, kingfishers, and more wildlife. After a brief walk to Lake Pilchicocha, a canoe takes you to the lodge, where you’ll settle in before a guided night walk or canoe ride to spot nocturnal species.

Day 3-4: Rainforest Ecolodge Exploration

Spend two days exploring the rainforest: walk canopy bridges for birdwatching, visit parrot-filled clay licks, paddle through swamps in search of caiman, anaconda, and capybara, or try piranha fishing. Back at the lodge, unwind with a lake swim or relax in your hammock.

Day 5: Returning to Quito 

After breakfast, take a canoe back to Coca for your flight to Quito. You’ll arrive in the early afternoon. From there, your onward journey continues, whether you’re returning home or exploring further.

Please note that we may visit sites in a different order.

Ships:

Set sail on an awe-inspiring expedition through the Galápagos Islands with Hurtigruten Expeditions. Aboard an eco-conscious expedition ship, you’ll experience the perfect blend of comfort and exploration as you get up close to the archipelago’s iconic wildlife, volcanic landscapes, and pristine shores. Below are the ships that bring this legendary journey to life.

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  • Antarctica Port Lockroy Ms Fridtjof Nansen Hx 43402 Photo Ted Gatlin
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Hx Hurtigrutenexpeditions Logo Rgb White 300dpi

Why Hurtigruten Expeditions?

HX offers over 130 years of adventure travel expertise, combining eco-friendly ships with immersive exploration of remote destinations. When booking through LaMacchia Travel, you’ll enjoy personalized service at no extra cost, ensuring a seamless and unforgettable journey.

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